There were four main model modifications (Ausführung A through D), which formed the basis for at least 22 variants. The initial idea was for a vehicle that could be used to transport a single squad of panzer-grenadiers to the battlefield protected from enemy small arms fire, and with some protection from artillery fire. In addition, the standard mounting of at least one MG 34 or MG 42 machine gun allowed the vehicle to provide support by fire for the infantry squad once they had disembarked in battle.
Positive aspects of the open top included greater situational
awareness and faster egress by the infantry, as well as the ability to
throw grenades and fire over the top of the fighting compartment as
necessary while remaining under good horizontal cover. The downside was a major vulnerability to all types of plunging fire; this included indirect fire from mortars and field artillery, as well as depressed-trajectory small arms fire from higher elevated positions, lobbed hand grenades, even Molotov's cocktails, and strafing by enemy aircraft.
The first two models were produced in small numbers from 1939. A and B models can be identified by the structure of the nose armor, which comprised two trapezoidal armor panels - the lower of which had a cooling hatch. The B model, which began production in 1940, eliminated the fighting compartment's side vision slits.
The C model, which started
production in mid-1940, featured a simplified hexagonal-shaped forward
armored plate for the engine. Models A through C had rear doors that
bulged out. The C model had a large production run, but was quite
complex to build, involving many angled plates that gave reasonable
protection from small arms fire. From early 1943, the D model was
developed with the purpose of halving the number of angled body plates,
simplifying the design and thus speeding up the production. D models can
be easily recognized by their single piece sloping rear (with flat
doors).
The standard personnel carrier version was equipped with a 7.92 mm
MG 34 or MG 42 machine gun mounted at the front of the open
compartment, above and behind the driver. A second machine gun could be
mounted at the rear on an anti-aircraft mount.
Variants were produced for specialized purposes, including with anti-aircraft guns, light howitzers, anti-tank guns and mortars or even large unguided artillery rockets, as well as a version with an infrared search light used to spot potential targets for associated Panther tanks equipped with infrared detectors.
Another potentially good design feature of the Sd.Kfz.251 was the
large track area, with the characteristic "slack track" design with no
return rollers for the upper run of track, and the Schachtellaufwerk
system of overlapping and interleaved main road wheels common to
virtually all German halftracks of the period.
This lowered the ground pressure
and provided better traction, giving the Sd.Kfz.251 better cross
country performance than most other nations' half-tracked vehicles.
However, the interleaved and overlapping main road wheels shared a major
problem with the Tiger I and Panther main battle tanks that also used such road wheel configurations - in muddy or winter weather conditions, such as those during a rasputitsa mud season or the coldest Russian winter conditions, accumulated mud and snow could freeze solid between the road wheels, immobilizing the vehicle.
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